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Kudela Raphael M. Kudela
Associate Professor of Ocean Sciences

Ocean Sciences Department
University of California
Santa Cruz, CA 95064
Office: A-461 Earth & Marine Sciences Bldg.
Tel: (831) 459-3290
Fax: (831) 459-4882

Email: kudela@ucsc.edu
Websites: KUDELA WEBSITE   |   LAB WEBSITE

Curriculum Vitae

Positions Held
Education
Professional Affiliations & Activities
Recent Research
Publications
Field Experience
Invited Talks
Sponsored Student Research
Courses Taught

POSITIONS HELD

1999-present Associate Professor, Ocean Sciences, University of California, Santa Cruz
2000-present Los Alamos National Laboratory Visiting Scientist
1999-present MBARI Collaborating Scientist
1998-present Associate Research Scientist, Romberg Tiburon Center, SFSU
1998-1999 Post-Doctoral Research Associate, Los Alamos National Laboratory
1996-1999 MBARI Post-Doctoral Fellow
1997-1998 Visiting Scientist, Romberg Tiburon Center, SFSU
1992-1995 NASA Global Change Fellow, USC
1992-1994 Research Consultant, MBARI
1991 Research Scientist, Oceans Division, Jet Propulsion Laboratory
1991 Research Consultant, The Cousteau Society

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EDUCATION

Ph.D., Biology (Biological Oceanography),
University of Southern California, 1995.

B.S. (Magna Cum Laude), Biology/Marine Science,
Drake University, 1989.

PROFESSIONAL AFFILIATIONS & ACTIVITIES

2005 Co-Chair, 52nd Annual Eastern Pacific Ocean Conference session “A Satellite Remote Sensing Primer for the New Millenium”, 27-30 September 2005, Stanford Sierra Camp, CA.
2004-2005 Member, National and Regional Organizing Committees for the US HAB Meeting (Asilomar, CA, 2005).
2004

Co-Chair, AGU Ocean Sciences session “Coastal Ocean Processes off the North American West Coast: Physical Dynamics and Biogeochemical Transformations IV”. January 26-30, Portland OR.

2002 Invited Chair, Ocean Optics XVI session “Ecology and Biochemistry II”, November 18-22, Santa Fe, NM
2001-Present Ocean Color Science Team Member (NASA)
1999-2000 Committee member and organizer, TRACER 2000 workshop (WS7: ASLO, Copenhagen; 2000)
1997-2000 SeaWiFS Science Team Member (NASA)
1996-Present Member, American Association for the Advancement of Science
1995-Present Member, International Society for Optical Engineering (SPIE)
1995-Present Member, Phycological Society of America
1994-Present Member, Committee on Space Research (COSPAR)
1992-Present Member, The Oceanography Society
1990-Present Member, American Society of Limnology and Oceanography
1990-Present Member, American Geophysical Union

RESEARCH PROJECTS

Overview:  Phytoplankton ecologist who wishes to understand the fundamental question: what controls phytoplankton growth and distribution in the ocean. More specifically, how do the multiple interactions of light, macro- and micronutrients and phytoplankton physiology determine the rates, processes, and patterns we observe in the marine environment? Oceanography is rapidly moving away from observational science towards an understanding of underlying mechanistic processes at all scales, in part because of the wealth of revolutionary new technological and scientific advances. My approach is to combine a suite of 3 tools: (1) remotely sensed data from moorings and satellites in combination with biological models; (2) novel bio-optical methods assaying phytoplankton physiology; and (3) the refinement of stable and radio-tracer isotopes.

Also see:    Kudela WebSite   and/or    Lab Page

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PUBLISHED WRITINGS AND CREATIVE ACTIVITIES   (Last five years)

Articles Submitted

Simmons, S.E., D.E. Crocker, R.M. Kudela, and D.P. Costa. Linking foraging behavior with bathymetry and oceanography in the sexually dimorphic northern elephant seal (Mirounga angustirostris). Marine Ecology Progress Series. Submitted July 2006.

Book Chapters

Bernard, S., R.M. Kudela, P. Franks, W. Fennel, A. Kemp, A. Fawcett, and G.C. Pitcher. 2006. The requirement for forecasting harmful algal blooms in the Benguela. In V. Shannon, G. Hempel, P. Malanotte-Rizzoli, C. Moloney and J. Woods (eds.). Large Marine Ecosystems, Vol. 14. Elsevier. [peer-reviewed]

Croll, D.A., and R. Kudela. Ecosystem impact of the decline of large whales in the North Pacific.  In J.A. Estes, D.P. DeMaster, D.F. Doak, T.M. Williams, and R.L. Brownell Jr. (eds.).  Whales, Whaling, and Ocean Ecosystems.  UC Press. [peer-reviewed]

Kudela, R.M. In press. Silicon:nitrogen interactions in the marine environment. In D. Bronk, M. Mulholland, D. Capone and E. Carpenter (eds). Nitrogen in the Marine Environment. Academic Press. [Invited chapter, peer-reviewed]

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Articles

Anderson, C.R., M.A. Brzezinski, L. Washburn, and R. Kudela. Accepted. Mesoscale circulation effects on a toxic diatom bloom in the Santa Barbara Channel, California. Marine Ecology Progress Series [peer-reviewed].

Armstrong, M.D., N. Ladizinsky, W.P. Cochlan, and R.M. Kudela. Accepted. Nitrogenous preference of toxigenic Pseudo-nitzschia australis (Bacillariophyceae) from field and laboratory experiments. Harmful Algae. [peer-reviewed]

Armstrong, M.D., and R. Kudela. Accepted. Evaluation of California isolates of Lingulodinium polyedrum for the production of yessotoxin. African Journal of Marine Science. [peer-reviewed]

Hickey, B., A. MacFadyen, W. Cochlan, R. Kudela, K. Bruland, and C. Trick. Accepted. Evolution of chemical, biological, and physical water properties in the northern California Current in 2005: Remote or local forcing? Geophysical Research Letters. [peer-reviewed]

Kudela, R.M., W.P. Cochlan, T.D. Peterson, and C.W. Trick. Impacts on phytoplankton biomass and productivity in the Pacific Northwest during the warm ocean conditions of 2005. Geophysical Research Letters. [peer-reviewed]

Kudela R.M., N. Garfield, and K.W. Bruland. Accepted. Bio-optical signatures and biogeochemistry from intense upwelling and relaxation in coastal California. Deep-Sea Research II. [peer-reviewed]

Largier J.L., C.A. Lawrence, M. Roughan, D.M. Kaplan, E.P. Dever, C.E. Dorman, R.M. Kudela, S.M. Bollens, F.P. Wilkerson, R.C. Dugdale, L.W. Botsford, N. Garfield, B. Kuebel-Cervantes, and D. Koracin. Accepted. WEST: a northern California study of the role of wind-driven transport in the productivity of coastal plankton communities. Deep-Sea Research II. [peer-reviewed]

Vander Woude A.J., J.L. Largier, and R.M. Kudela. Accepted. Nearshore retention of upwelled waters north and south of Point Reyes (northern California) – patterns of surface temperature and chlorophyll observed in CoOP WEST. Deep-Sea Research II. [peer-reviewed]

Weise, M.J., D.P. Costa, and R.M. Kudela. Accepted. At-sea movement and diving behavior of male California sea lion (Zalophus californianus) during 2004 and 2005. Geophysical Research Letters. [peer-reviewed]

Dierssen, H.M., R.M. Kudela, J.P. Ryan, and R.C. Zimmerman. 2006. Red and black tides: Quantitative analysis of water-leaving radiance and perceived color for phytoplankton, colored dissolved organic matter, and suspended sediments. Limnology and Oceanography, 51(6): 000-000 (in press). [peer-reviewed]

Kudela, R., G. Pitcher, T. Probyn, F. Figueiras, T. Moita, and V. Trainer. 2005. Harmful Algal Blooms in coastal upwelling systems. Oceanography 18(2): 184-197. [peer-reviewed]

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Ryan, JP, Dierssen, HM, Kudela, RM, Scholin, CA, Johnson, KS, Sullivan, JM, Fischer, AM, Rienecker, EV, McEnaney, PR, and Chavez, FP. 2005. Coastal ocean physics and red tides: An example from Monterey Bay, California. Oceanography 18(2): 214-223. [peer-reviewed]

Kudela, R.M., W.P. Cochlan, and A. Roberts. 2004. Spatial and temporal patterns of Pseudo-nitzschia spp. in central California related to regional oceanography. Harmful Algal Blooms 2002. Steidinger, K. et al. (Eds.). Intergovernmental Oceanographic Commission of UNESCO. [peer-reviewed]

Kudela, R.M., A. Roberts, and M. Armstrong. 2004. Laboratory Analyses of Nutrient Stress and Toxin Production in Pseudo-nitzschia spp. from Monterey Bay, California. Harmful Algal Blooms 2002. Steidinger, K. et al. (Eds.). Intergovernmental Oceanographic Commission of UNESCO. [peer-reviewed]

Kudela, R.M. and F.P. Chavez. 2004. The impact of coastal runoff on ocean color during an El Niño year in central California. Deep-Sea Research II, in press. doi:10.1016/j.dsr2.2004.04.002. [peer-reviewed]

Coale, K.H., Johnson, K.S., Chavez, F.P., Buesseler, K.O., Barber, R.T., Brzezinski, M.A., Cochlan, W.P., Millero, F.J., Falkowski, P.G., Bauer, J.E., Wanninkof R.H., Kudela, R.M., Altabet, M.A., Hales, B.E., Takahashi, T., Landry, M.R., Bidigare, R.R., Wang, X., Chase, Z., Strutton, P.G., Friederich, G.E., Gorbunov, M.Y., Lance, V.P., Hilting, A.K., Hiscock, M.R., Demarest, M., Hiscock, W.T., Sullivan, K.F., Tanner, S.J., Gordon, R.M., Hunter, C.N., Elrod, V.A., Fitzwater, S.E., Jones, J.L., Tozzi, S., Koblizek, M., Roberts, A.E., Herndon, J., Brewster, J., Ladizinsky, N., Smith, G., Cooper, D., Sheridan, C.C., Twining, B.S., Johnson, Z.I.  2004. Southern Ocean Iron Enrichment Experiment: Carbon cycling in high- and low-Si waters. Science 304:408-414. [peer-reviewed]

Berelson, W., J. McManus, K. Coale, K. Johnson, D. Burdige, T. Kilgore, D. Colodner F. Chavez, R. Kudela, J. Boucher. 2003. A time series of benthic flux measurements from Monterey Bay, CA. Continental Shelf Research, 23: 457-481. [peer-reviewed]

Kudela, R.M. and F.P. Chavez. 2002. Multi-platform remote sensing of new production in central California during the 1997-1998 El Niño.
Progress in Oceanography 54: 233-249.. [peer-reviewed]

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Other

Pitcher, G., F. Figueiras, R. Kudela, T. Probyn, T. Moita, S. Sanchez, V. Trainer, E. Urban, and H. Enevoldsen. 2006. GEOHAB Core Research Project—HABs in upwelling systems: call for participants. Harmful Algae News, 31: 9-11.

Pitcher, G., T. Moita, V. Trainer, R. Kudela, P. Figueiras, and T. Probyn. 2005. GEOHAB Core Research Project: HABs in upwelling systems. SCOR and IOC, Baltimore and Paris.

Prevention, Control, and Mitigation of Harmful Algal Blooms: A Research Plan. 2001. Submitted to the United States Congress by the National Sea Grant Program.

Published Extended Abstracts

Kudela, R.M. 2006. Linking the physiology and ecology of the harmful red tide organism Cochlodinium to better understand bloom events. Recent progress on the research and management of Cochlodinium blooms Extended Abstracts, Cheju National University, Republic of Korea.

Cochlan, W.P. and R.M. Kudela. 2004. The Southern Ocean Iron Enrichment Experiment: The nitrogen uptake response and potential effects of ammonium inhibition.  PICES Scientific Report: Proceedings of the IFEP Workshop.

Ryan, J., F. Chavez, J. Bellingham, E. Rienecker, H. Dierssen, R. Kudela, A. Vander Woude, and R. Maffione. 2002. Environmental processes in the Monterey Bay National Marine Sanctuary: Studies integrating AVIRIS and synoptic in situ sensing. Proceedings of the 2004 JPL Airborne Earth Science Workshop. NASA, Jet Propulsion Laboratory, Pasadena, California, USA.. JPL Publication 87-38.

Ryan, J., F. Chavez, J. Bellingham, E. Rienecker, H. Dierssen, R. Kudela, A. Vander Woude, and R. Maffione. 2002. Environmental processes in the Monterey Bay National Marine Sanctuary: Studies integrating AVIRIS and synoptic in situ sensing. Proceedings of the 2004 JPL Airborne Earth Science Workshop. NASA, Jet Propulsion Laboratory, Pasadena, California, USA.. JPL Publication 87-38.

Kudela, R.M., N. Garfield, L. Kaupp, G. Smith, and K.W. Bruland. 2002. Bio-optical signatures from intense upwelling and relaxation in coastal California. SPIE, Ocean Optics XVI (CD-ROM distribution).

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FIELD EXPERIENCE (Major field programs)

September 2006 Coastal Ocean Applications and Science Team (COAST, NOAA: R/V John Martin)
June, August 2005 River Influences on Shelf Ecosystems (NSF CoOP; R/V Wecoma)
July 2004 River Influences on Shelf Ecosystems (NSF CoOP; R/V Wecoma)
August-Sept 2003 Bering Sea Cruise (R/V Kilo Moana, UCSC; collaboration with Dr. K. Bruland)
August 2002 Shelf-Break Krill Study (R/V Zephyr, MBARI; collaboration with Dr. J. Ryan)
July 2002 Wind to Whales (monthly cruises, Monterey Bay; R/V John Martin)
June 2002 Chief Scientist, Wind-Driven Ecosystems and Shelf Transport (NSF CoOP; R/V Point Sur)
May-June 2001 Wind-Driven Ecosystems and Shelf Transport (NSF CoOP; R/V Point Sur)
September 2000 Class cruise, Ocea 130/230, R/V Point Sur
May-June 2000 Wind-Driven Ecosystems and Shelf Transport (NSF CoOP; R/V Point Sur)
March 1999 NOAA/NMFS Cruise (in support of NASA grant # NAG5-7632)
1997-1999 Studies of Ecological and Chemical Responses to Environmental Trends (SECRET)
  R/Vs New Horizon, Point Sur, Western Flyer, Central California (8 cruises)
May-June 1995 Iron-Ex II,  R/V Melville, Equatorial Pacific
April 1995 Coastal Ocean Processes (CoOP),  R/V Point Sur, Central California
March 1995 Orange Coast College  R/V Marda, Newport Beach, California
March 1994 World Ocean Circulation Experiment (WOCE), NOAA ship R/V Discoverer, Eq. Pacific
1992-1993 Shift-Up Program R/V Point Sur, Central California (6 cruises)
1990-1991 Flow Around Abrupt Changes in Topography (TOPO)
  R/Vs Thomas G. Thompson, New Horizon, western Pacific (2 cruises)
1990-1991 Monterey Bay Food Web Scavenging Project  R/V Point Sur (9 cruises)
April-May 1990 EPOCS  NOAA ship R/V Malcolm Baldridge, Equatorial Pacific
May 1989 Search for Cold Water  R/V Pt Lobos, Monterey Bay

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INVITED SCIENTIFIC PRESENTATIONS

“Linking the physiology and ecology of the harmful red tide organism Cochlodinium to better understand bloom events”, Recent progress on the research and management of Cochlodinium blooms workshop, 22-24 May, 2006, Cheju National University, Republic of Korea.

“Harmful Algal Blooms in upwelling systems: physical-biological coupling underlying HABs”. International Workshop on Forecasting and Data Assimilation in the Benguela and Comparable Systems, Cape Town, 8-11 November 2004.

“Light-nutrient interactions in a not so steady ocean” Carnegie Institute for Global Ecology Seminar Series December 2003.

“Lessons learned from the harmful diatom Pseudo-nitzschia—Can we really predict what will happen in the ocean?”  Stanford Seminar Series November 2003.

"Modeling the Biological Impacts of El Niños in Central California". Water Resources Seminar Series, USGS Menlo Park, April 2002.

“Estimating the Impact of El Niños / La Niñas in Central California Using Remote Sensing”. University of California Santa Cruz, Remote Sensing Seminar Series, Winter 2002.

“Silicon, Nitrogen, Iron and Light: What Really Controls Diatom Production?” Moss Landing Marine Labs public lecture, January 2002.

“Silicon Limitation of Phytoplankton: More Important Than You Might Think”, Monterey Bay Aquarium Research Institute, November 1, 2000.

Public Lecture or Forum Participation

“Do Pristine Coastal Waters Exist? The Role of Naturally Occurring Toxins in California”, 2006 Spring Symposium, Northern California Society of Toxicology, 28 April 2006, Half Moon Bay CA.

Invited speaker, Inaugural Lecture Series at Seymour Center, Long Marine Lab. “Phytoplankton Dynamics – The Grass of the Sea”, April 18, 2000.

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SPONSORED STUDENT & POSTDOCTURAL RESEARCH

Post-Doctoral Scholars

2005-Present Tawnya D. Peterson
2000-2004 Bethany Jenkins  (co-advised with Jon Zehr)
  Current Position: Assistant Professor Department of Cell and Molecular Biology and Graduate School of Oceanography

Past Graduate Students

M.S. (2003) Demetres Koutroukides
  Dr. Earl H. and Ethel M. Myers Oceanography and Marine Biology Trust Fellowship recipient (2001)
  Thesis Title: “Modeling chlorophyll in San Francisco and Bodega Bays Using SeaWiFS”

Current Graduate Students
(Advanced to Candidacy)

Admitted Fall 2001 Meredith Armstrong
Fall 2005
Advanced to Candidacy
2005
Marilyn C. Davis Memorial Scholarship,  Coastal Environmental Quality Initiative Graduate Student Fellowship
2004

Outstanding Student Achievement Award, Ida Benson Lynn Fellowship, Sea Grant Rapid Response Grant

2002, 2003

Ocean Science Outstanding Teaching Assistant Nominee, Ocean Science Research and Travel Award

2001

Earl and Ethyl Myers Trust Grant, STARS Scholarship

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Ph.D. and M.S. Students

Admitted Fall 2001 Shelly Benoit, Ph.D. student
 (summer 2003)
EU Surface Ocean Lower Atmosphere Study (SOLAS) Summer School participant, Italy
   
Admitted Fall 2006 Melissa (Misty) Blakely, M.S. student
 

 

Admitted Fall 2006

Jenny Quay Lane

 

CDELSI Graduate Fellowship recipient (2006)

 

 

Admitted Fall 2003

Sherry Palacios, Ph.D. student

 

 

Admitted Fall 2006

Anna Rubin, M.S. student

Other - Graduate Student Thesis Committees

Sibel Bargu Ph.D. in Ocean Sciences (completed 2001)
Carolyn Berger  M.S. in Ocean Sciences
Kristen Buck Ph.D. in Ocean Sciences
Christy Caldwell M.S. in Ocean Sciences
Olivia Cheriton  Ph.D. in Ocean Sciences
Itchung Cheung M.S. in Ocean Sciences
Aaron Cozen Ph.D. in Ocean Sciences
Eric Danner Ph.D. in Ecology and Evolutionary Biology (completed 2006)
Laura Lessin M.S. in Ocean Sciences (completed 2002)
Amy Little  M.S. in Ocean Sciences (completed 2003)
Allison Luengen Ph.D. in Environmental Toxicology
Erin McCarthy M.S. in Ocean Sciences
Anna Pfeiffer-Hoyt M.S. in Ocean Sciences (completed in 2005)
Daria Siciliano Ph.D. in Ecology and Evolutionary Biology (completed in 2005)
Samantha Simmons M.S. in Ecology and Evolutionary Biology (completed 2003)
Samantha Simmons Ph.D. in Ecology and Evolutionary Biology
Veronica Vigilant Ph.D. in Ocean Sciences
Andreas Walli Ph.D. in Ecology and Evolutionary Biology
Shaye Wolf M.S. in Ocean Sciences (completed 2002)
Shaye Wolf Ph.D. in Ecology and Evolutionary Biology
Victoria Welborn M.S. in Ocean Sciences
Michael Wiese Ph.D. in Ecology and Evolutionary Biology (completed 2006)
Clarissa Anderson Ph.D. in Ecology, Evolution, and Marine Biology (UC Santa Barbara)
Erich Reinecker M.S. in Oceanography (Moss Landing Marine Laboratory)

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COURSES TAUGHT (At University of California, Santa Cruz)

Course Number Course Name
OCEA 101 Marine Environment
OCEA 130/230+BIOL 159 Biological Oceanography
OCEA 290H
Topics in Ocean Optics
OCEA 292 Ocean Sciences Seminar
OCEA 293 Graduate Research Seminar
OCEA 299
Thesis Research

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